Table of Contents

Volume 36, 2026
Articles
Médail F.
Vol. 36 | pp. 5-17

This work is a contribution to the knowledge of the vascular flora of the archipelago (SE Greece), which is part of the South Anatolian biogeographical province. An autumn survey (October 2025) identified nine species new for the archipelago (Amaranthus viridis, Dittrichia graveolens, Eclipta prostrata, Euphorbia prostrata, Gastridium phleoides, Orobanche minor, Portulaca oleracea, Setaria adhaerens, Sorghum halepense) and clarified the distribution of several taxa of great biogeographical interest (e.g. Campanula kastellorizana, Colchicum macrophyllum, Cyclamen maritimum, Dianthus muglensis, Polygonum praelongum, Thliphthisa brevifolia). A checklist is also presented for Aghios Georgios, a small satellite islet (1 ha) with a high species richness as 149 plant taxa have been recorded in total. The Kastellorizo archipelago, rich in about twenty endemic vascular plants, is a very interesting biogeographical entity that deserves increased protection of its biodiversity.

Bougaham A.F.
,
Bencheikh R.
,
Bouchareb A.
,
Sahnoune M.
Vol. 36 | pp. 19-29

Euonymus europaeus (Celastraceae) is a deciduous forest tree species of Eurasian origin, widely distributed across Europe and western Asia, but hitherto unrecorded in North Africa. Recent botanical surveys carried out in the Babors Mountains (northeastern Algeria) have confirmed the presence of this species in three isolated mixed deciduous forests located between 700 and 1100 m a.s.l., including the Taza National Park and the forests of Tloudène and Tazegzeout. The identification is based on diagnostic morphological characters, particularly the non-winged capsules and the quadrangular young branches. Comparison with Euonymus latifolius var. kabylicus, the only taxon of the genus previously reported from Algeria, clearly supports the attribution of the studied populations to Euonymus europaeus. The restricted distribution of the species within relict montane forests, together with its absence from North African floristic literature, suggests an ancient native status, likely related to the role of the Babors Mountains as biogeographical refugia. This finding highlights the floristic and conservation value of the region and underlines the need for further botanical investigations in northeastern Algeria.

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Résumé

Euonymus europaeus (Celastraceae) est une espèce arborée forestière à feuillage caduc, d'origine eurasiatique, largement répandue en Europe et en Asie occidentale, mais jusqu'à présent non signalée en Afrique du Nord. Des inventaires botaniques...

Gjeta E.
,
Damo R.
,
Barina Z.
,
Icka P.
Vol. 36 | pp. 31-36

This contribution updates the distribution of three vascular plant taxa in various habitats and substrates within the Korça region in Southeastern Albania. Two of these: Trigonella graeca and Aremonia agrimonoides subsp. pouzarii are native to Albania, while Trigonella caerulea is a casual alien species likely introduced as an aromatic plant, many years ago with T. foenum– graecum. Both A. agrimonoides subsp. pouzarii and T. caerulea are reported here as new records for Albania; the former is a Balkan taxon and the latter is a cultigen originating from the Eastern Mediterranean, Mediterranean Europe, and Asia. Furthermore, a new Albanian southeastern locality and three southern unpublished ones are given for T. graeca, representing a significant expansion to the distribution range both within the country and across the Balkan Peninsula.

Matevski V.
,
Tan K.
,
Kostadinovski M.
,
Ćušterevska R.
Vol. 36 | pp. 37-44

This paper discusses some aspects concerning the year of publication and the locus classicus of Senecio macedonicus Griseb. New data on the distribution of this species within the territory of the Republic of North Macedonia are given.

Boulemtafes A.
,
Saadi A.
Vol. 36 | pp. 45-48

This article documents the first record of Petunia ×atkinsiana escaped from ornamental cultivation in Algeria. This South American nothospecies was observed in the commune of Seraidi (Annaba Province). We classify P. ×atkinsiana as casual alien showing neither substantial spread nor clear signs of naturalization.

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Résumé

Dans le présent article est documentée le premier signalement de Petunia ×atkinsiana échappée de culture ornementale sur le territoire algérien. Cette nothoespèce sud-américaine a été observée dans la commune de Seraidi (Wilaya de Annaba). Nous classons P. ×atkinsiana comme ayant un statut d’exotique occasionnelle ne montrant ni propagation substantielle ni signes clairs de naturalisation.

El Mokni R.
,
El Mokni A.
,
El Mokni R.
Vol. 36 | pp. 49-62

During botanical surveys undertaken from northern to southern Tunisian territory, mostly within the last two decades, new occurrences of alien vascular plants (lianas) were detected. These records include: Campsis grandiflora, Lablab purpureus subsp. uncinatus and Tinospora cordifolia new to the Mediterranean area; Solandra maxima new to the African inland and Ipomoea pes-caprae new to North Africa. In addition, new national distributional data for Ipomoea batatas, Lablab purpureus subsp. purpureus and Momordica balsamina, already known from Tunisia, are here reported. All taxa are considered casual or naturalized aliens where the horticultural trade remains the most likely pathway for their introduction. General macromorphological characteristics together with original photographs are provided. Actual distribution, ecological notes (habitat, phenology) as well as the degree of naturalization in Tunisia are presented.

Calvia G.
,
Fois M.
,
Mascia F.
,
Podda L.
,
Manca M.
,
Bertotto G.
,
Campus G.
,
Lallai A.
,
Ruggero A.
,
Porceddu M.
,
Rivieccio G.
,
Cuena-Lombraña A.
,
Bagella S.
,
Farris E.
,
Bacchetta G.
Vol. 36 | pp. 63-98

Following the first two contributions for a new, updated, and revised vascular flora of Sardinia, new data about the distribution of 30 taxa growing in the island are here presented. Taxa are both native and alien, and are characterized by their very low frequency, phytogeographical significance, endemicity, conservation status, invasiveness or novelty to Sardinia and Italy. The results derive from floristic research carried out throughout Sardinia and various neighbouring islands and islets. The first reports of Bulbine frutescens, Coleus barbatus, Dasylirion serratifolium, Euphorbia ingens, and E. milii are here recorded for the first time in Italy (casual alien). Additionally, Ageratum conyzoides, Cyperus papyrus, Erysimum odoratum, Galinsoga quadriradiata, and Narcissus trumpet daffodil group (casual alien taxa) are documented here for the first time in Sardinia. Moreover, the confirmations for Sardinian flora of Carex acutiformis, Catananche lutea, and Suaeda splendens are here provided after 40 years without new reports. New distributional data for 12 taxa, most of which known for their rarity, are gathered. Notably, Callitriche lusitanica was previously reported only for a few sites. Concerning other taxa, a clarification of their distribution range on the island is documented, as for the rare Ceratophyllum submersum subsp. submersum, Limoniastrum monopetalum, and Malva stenopetala subsp. plazzae, which have recently faced contract...

Bouchet M.
,
Molina J.
,
Martínez Labarga J.M.
Vol. 36 | pp. 99-106

Linum corymbiferum, a species treated as historic casual in France, was rediscovered in the department Gard, on its previously documented location and several close places. This species, endemic to Algeria and Tunisia, was discovered in 1875 in Rodilhan (department Gard, France) by A. Barrandon, J. Duval-Jouve and Courcière. It was then recorded four times at the same location in the following years by different botanists who collected samples. Other herbarium specimens dated 1872, 1913 and 1927 were respectively collected in Hyères (Var department, France) and Béziers (department Hérault, France). Subsequently, the species had not been recorded in Europe until 2025.

Iamonico D.
,
Troìa A.
Vol. 36 | pp. 107-111

The name Rosa viscosa (Rosaceae), validly published by Gussone in his Supplementum ad Florae Siculae Prodromum, was recently lectotypified by Iamonico & Lattanzi on a specimen included in Jan’s collection and preserved at the Herbarium NAP. These authors provided an erroneous indication of the original reference of the name, citing Jan’s Catalogus Plantarum Phaenogamarum, ad Usum Botanophilorum Exsiccatarum (published in 1818) instead of Gussone’s Supplementum ad Florae Siculae Prodromus (published in 1832). Actually, the name R. viscosa does not appear in Jan’s Catalogus, while we traced it into a later published catalogue, i.e. Jan’s Elenchum Plantarum (published in 1831). However, R. viscosa is there invalid being a nomen nudum. So, since the name reported in the taxonomic treatment by Iamonico & Lattanzi is linked to a reference (Jan’s Catalogus Plantarum Phaenogamarum) where that name does not occur, the typification would not be applicable, and a revision is here proposed. It should be noted that a clear indication in the Madrid Code as to how to handle a situation like this does not appear. Furthermore, we noted that Jan’s catalogue is reported as original reference of R. viscosa by all the major databases of plant names and other sources, whereas Jan’ Elenchus is never cited.

Chatelain C.
,
Chambouleyron M.
,
Mombrial F.
,
Léger J.-.
,
Perret M.
Vol. 36 | pp. 113-128

The rediscovery of two little-known species, Foeniculum subinodorum and Anethum theurkauffii in Morocco and Mauritania has led us to revise the taxonomy of the genus Anethum, with support of molecular data (nrDNA ITS and plastid rps16 sequences). Our phylogenetic analyses and morphological observations confirm that these two taxa are the same species to be renamed under Anethum subinodorum. Moreover, the phylogenetic analysis of the whole genus Anethum led us to recognize the close relationship between A. subinodorum and A. foeniculoides; and confirm that Foeniculum scoparium from Chad should be treated as a synonym of A. foeniculum. We recognize eight taxa (seven species and one subspecies) in Anethum, for which we propose an identification key and finally we here propose one new combination and design two lectotypes . These results confirm that Morocco is the center of diversity of Anethum with two strict sub-Saharan endemic species and a Mediterranean one.

Tavilla G.
,
Lanfranco S.
Vol. 36 | pp. 129-138

The name Melitella pusilla, basionym for the accepted Crepis pusilla, is lectotypified. The name was originally published within the monotypic new genus Melitella basing on plants collected by Sommier on Gozo Island (Maltese Archipelago). Specimens preserved at E, FI, GH, K, LY, MU, MW, VT, P, PI, SLA, VAL, WU, and Z herbaria have been examined. The lectotype is a specimen at FI. An earlier potential typification is dismissed, as the corresponding herbarium sheet bears plants collected on two different dates and is not therefore one specimen. Furthermore, M. pusilla var. laciniata, lacking any traceable original material, is neotypified with a specimen kept at VAL.

Akhrif I.
,
Ismaili S.E.
,
Maurady A.
,
Britel M.R.
,
Homrani Bakali A.
Vol. 36 | pp. 139-156

This study examines the physical properties of seeds from seven species of Stachys (S. arenaria, S. circinata, S. fontqueri, S. germanica, S. officinalis, S. saxicola, and S. mouretii) to assess their morphological diversity and potential applications. Measurements included seed dimensions, derived geometric parameters, density, porosity, moisture content, and thousand-seed mass. Results revealed significant variation across taxa. S. germanica had the largest seeds, the highest bulk density (601.77 kg m−3), a high true density (991.95 kg m−3), and the lowest porosity (38.78%), while S. officinalis showed the highest moisture content (13.81%) and a notably low aspect ratio (0.56). These findings provide quantitative seed-morphological data useful for species comparison, seed handling, and future integrative studies.

Conti F.
,
Falcinelli F.
,
Palermo D.
,
Giacanelli V.
,
Koopman J.
,
Więclaw H.
,
Bartolucci F.
Vol. 36 | pp. 157-168

New or revised plant records of 42 taxa (species and subspecies) are reported for Italy, of which 25 are natives and 17 aliens. Hypericum richerii subsp. grisebachii is new for Italy, Dianthus carthusianorum subsp. atrorubens is new for central Italy, and another 12 natives and aliens are new or confirmed for Umbria, nine for Abruzzo, 11 for Molise, four for Lazio and one for Marche, Basilicata, Campania and Calabria. Eight taxa are to be excluded from the flora of Abruzzo, and one from Umbria, Marche, Lazio, Molise, Campania and Basilicata.

Tanji A.
,
Khamar H.
Vol. 36 | pp. 169-179

During weed surveys in irrigated crops in the Kenitra province of Morocco in 2024 and 2025, we found three new Poaceae weeds: Dinebra panicea subsp. mucronata, Diplachne fusca subsp. fascicularis, and Oryza punctata. At present, the three semi-aquatic taxa are naturalized and adapted to water-saturated conditions prevailing in rice and sugarcane fields. To facilitate their identification, this note provides a brief morphological description of each taxon, photographs, a distribution map, and data on ecology and habitats. These taxa are considered new alien records for the flora of Morocco.

Yüceol F.
Vol. 36 | pp. 181-190

Cucumis maderaspatanus (Cucurbitaceae) is reported here as a new species for the flora of Türkiye. Specimens of this species, which is native to tropical Africa and South Asia, were collected from a field margin and a roadside in a disturbed habitat in the Tarsus district of Mersin Province in southern Türkiye. The identity of the species was confirmed by detailed morphological examination of leaves, flowers and fruits under the stereomicroscope. In this study, a comprehensive description of the species, detailed photographs and a distribution map are provided. This record has been added to the Turkish flora as a new casual alien species.

Zediri H.
,
Belabed A.I.
,
Hamel T.
Vol. 36 | pp. 191-213 | Suppl. material

The Guerbes-Senhadja wetland complex (wilaya of Skikda, north-eastern Algeria) is one of the most floristically significant wetland systems of Numidia, yet the ecology of its benthic macrophyte communities has remained largely undocumented. This study presents the first systematic inventory of aquatic macrophytes across nine representative waterbodies of the complex, conducted during the spring of 2025, alongside an analysis of the physicochemical parameters structuring their distribution. A total of 31 benthic macrophyte taxa belonging to 12 families and 16 genera were recorded. Floristic richness varied markedly with habitat type: saline semi-temporary marshes harboured four to seven taxa, temporary ponds supported up to 13 species, while peaty lakes and alder carrs yielded 11 to 18 taxa with greater abundance. The broadly distributed chorological element dominated (38.71%), followed by the northern (29.03%) and Mediterranean (25.80%) elements. Seventeen taxa were identified as holding patrimonial value, including 14 rare or very rare species, 7 protected under Algerian law, and 9 listed on the IUCN Red List. Six new floristic records for western Numidia are reported. A Principal Component Analysis (PCA) of 18 environmental and physico-chemical variables distinguished three ecological site typologies along gradients of anthropogenic disturbance, topography and hydrological regime. A co-inertia analysis (RV = 0.63, p < 0.05) confirmed a significant co-structure between the m...

Seregin A.P.
,
Yevseyenkov P.E.
,
Svirin S.A.
,
Kashirina E.S.
Vol. 36 | pp. 215-238

This paper summarises data on the flora of Sevastopol area (Crimean Peninsula) accumulated between 2015 and 2026. The paper presents records of 104 new plant species identified within the area, including three confirmations of historical records from literature. Of these, 69 species are classified as native, while 35 are alien plants at various stages of naturalisation. Within this list, four species were reported for the first time in Eastern Europe when discovered (Gelasia villosa, Lomelosia brachiata, L. divaricata, and Aegilops peregrina), and a further nine species were new records for the Crimean Peninsula (Buglossoides incrassata, Iris halophila, Ludwigia brevipes, Chloris virgata, Polypogon maritimus, Potamogeton sarmaticus, Ranunculus circinatus, Sicyos angulatus, and Thinopyrum acutum). Consequently, the flora of the Sevastopol area now comprises 1,745 species of vascular plants (excluding literature records not supported by herbarium specimens or recent photographic evidence).

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FLORA MEDITERRANEA
Published on behalf of the International Foundation pro Herbario Mediterraneo by
Francesco M. Raimondo & Werner Greuter
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Gianniantonio Domina
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Francesco M. Raimondo & Werner. Greuter
Editorial Committee
Svetlana Bancheva Nikolova (Bulgarian Academy of Sciences, Sofia), Cesar Blanché (University of Barcelona, Barcelona), Patrizia Campisi (University of Palermo, Palermo), Gianniantonio Domina (University of Palermo, Palermo), Eleonora Gabrielian (National Academy of Sciences of Republic of Armenia, Erevan, retired), Fabio Garbari (University of Pisa, Pisa, retired), Werner Greuter (Freie University of Berlin, Berlin, Emeritus), Stephen L. Jury (University of Reading, Reading, retired), Georgia Kamari (University of Patras, Patras, Emeritus), Giuseppe Antonio Malfa (University of Catania, Catania), Natalizia Miceli (University of Messina, Messina), George Nakhutsrishvili (Ilia State University, Tbilisi, retired),  Francesco M. Raimondo (University of Palermo, Palermo, retired), Cristina Salmeri (University of Palermo, Palermo), Sonja Siljak-Yakovlev ( University Paris-Sud, Orsay), Vivienne Spadaro (University of Palermo, Palermo), Benito Valdés (University of Sevilla, Sevilla, retired) & Giuseppe Venturella (University of Palermo, Palermo).
Referees Committee
Pepy Bareka (Agricultural University of Athens, Athens), Attilio Carapezza (University of Palermo, Palermo, retired), Régis Courtecuisse (University of Lille, Lille), Vincent Demoulin (University of Liège, Liège, honorary),  Mathias Erben (University of Munchen, Munchen, retired), Mohamed Fennane (University Mohamed V, Rabat), Adil Güner (University of Istanbul, Istanbul, retired), Daniel Jeanmonod (Conservatoire et Jardin botaniques de la Ville de Genève, Genève, honorary curator), Ekaterina Kozhuharova (Medical University of Sofia, Sofia), Anna M. Mannino (University of Palermo, Palermo), Marta Mariotti Lippi (University of Firenze, Firenze, retired),  Frédéric Médail (Aix Marseille University,Marseille), Enio Nardi (University of Firenze, Firenze, retired), Fatima N. Özhatay (University of Istanbul, Istanbul, retired), Gonzalo Nieto Feliner (CSIC, Madrid), Pier Luigi Nimis (University of Trieste, Trieste), Alessio Papini (University of Firenze, Firenze), Lorenzo Peruzzi (University of Pisa, Pisa), Dimitri Phitos (University of Patras, Patras,  Emeritus), Rosa M. Ros Espín (University of Murcia, Murcia), Federico Selvi (University of Firenze, Firenze), Shalva Sikharulidze (Ilia State University, Tbilisi),  Arne Strid (University of Copenhagen, Copenhagen, retired), Errol Véla (University of Montpellier, Montpellier) & Brigitte Zimmer (Freie University of Berlin, Berlin, retired).
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